Auto-rejected from a great match, so I found a way to follow up...

The hiring staff replied that I was missing CSS as a qualification. Now, I have 12 years of frontend work on my resume. But it turns out, upon review, that I wrote "HTML/CSS" in my skills junk drawer section. Moral is, no matter how good your bullets are, make your keywords space delimited. Your first audience is a RegEx. Also if something feels off, follow up. Might take some digging to find the right channel, but be polite and not much can go wrong. Update: for posterity, I ended up getting a great offer for this position and took it. I plan on mentioning the issues with their screening after I start there.